Naturalis is official partner in the BSc and MSc Biology curriculums of Leiden University, Wageningen University, the University of Amsterdam, and the University of Applied Sciences Leiden. Naturalis staff also participates in BSc and MSc teaching in Earth Sciences (Utrecht University, Free University of Amsterdam), and the BSc curriculum of Sciences (University of Maastricht). Naturalis offers courses at BSc, MSc and PhD level, and supervises term papers and BSc, MSc and PhD research projects. The subjects offered include systematics, phylogenetics, biogeography, palaeontology, genomics, bioinformatics, marine biology, evolutionary biology, taxonomy of selected groups (arthropods, molluscs, land plants, and fungi).