- Systematics and evolution of Cruciferae
- Floristics of Israel and Jordan
- Arachnid systematics and taxonomy, comparative morphology, phylogeny; Myriapoda, Chilopoda, Diplopoda, Arachnida, Araneae, Scorpiones, Pseudoscorpiones, Opiliones, Solifugae, Acari, Amblypygi, Palpigradi
- Mollusc of Israel and neighbouring areas.
- Reptiles and amphibians of Israel and neighbouring areas.
- Fish of Israel and neighbouring areas
- Vertebrate palaeontology of Mio-Holocene of the southern Levant
- Conservation genetics of endangered species
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