- Mammals, reptiles and amphibians of the eastern Mediterranean area
- Conservation of islands
- Biodiversity - biology - ecology - conservation of beetles of Balkans and the eastern Mediterranean area
- Biodiversity - biology - ecology - conservation - biogeography of terrestrial molluscs
- Taxonomy of Chilopoda of the Eastern Mediterranean and the European region
- Biodiversity - Ecology - Biology and Conservation of bats
- Taxonomy - Ecology and Phylogeography of scorpions of the Mediterranean area
- Ecology of aquatic invertebrates (all groups) of the eastern Mediterranean area
- Cretan and eastern Mediterranean flora
- Molecular taxonomy
- Distribution – Ecology – Conservation of birds of the eastern Mediterranean area
- Management of wetlands of the eastern Mediterranean area
- Structural Geology, Geomorphology, Geodiversity, Geoconservation
- Hydrogeology
- Terrestrial vertebrate and marine invertebrate palaeontology
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