- Palaeontology: fossil amphibians and reptiles, mammals (focus insectivors), fossil insects (focus dragonflies), aquatic vertebrates, Cenozoic molluscs, Mesozoic ammonites and crustaceans,
- Mesozoic and Cenozoic red algae (Corallinaceae), Palaeogene and Neogene phanerogams, plant physiology
- Botany: cryptogams (mainly mosses), phanerogams
- Zoology: mammals, birds, fish, molluscs, amphibians and reptiles
- Entomology: Ephemeroptera, Diptera: Tachinidae, Hymenoptera: Chalcidoidea, Coleoptera:
- Tenebrionidae, Lepidoptera: Bombycoidea, Geometridae, Sesiidae, Araneae: Salticidae
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