- Zoology Unit: Arthropoda, Insecta: Lepidoptera, Elachistidae, Coleoptera, Elateriformia, Dytiscidae and Chrysomelidae, Macroplea, Diptera, Syrphidae and Platypezidae, Sciaridae, Hymenoptera, Chrysididae
- Crustacea, Mysida and Amphipoda, Araneae, Platyhelminthes, Cestoda
- Botany Unit: Vascular plants, Chenopodiaceae, Salicornia, Musa and Cyperus sect. Arenarii, Oxandra, Asteraceae and Cotoneaster, Symphytum
- Hepatic systematics, phytogeopgraphy, Western Melanesian hepatics, Chinese hepatics, epiphytes, Helsinki bryophytes, medicinal plants
- Taxonomy, systematics and evolutionary history of bryophytes, especially Lejeuneaceae and Schistochilaceae
- Phytogeography, taxonomy and classification of bryophytes, Mniaceae, Philonotis, Rhodobryum, bryoflora of Finland, China and New Guinea
- Molecular phylogeny of liverworts
- Sphagnum. History of bryology, nomenclature of bryophytes
- Mycology, Ascomycetes, lichen forming fungi in particular, Evolution of the Basidiomycetes, Lichen genus, Lichenicolous fungi growing on Bryoria, Lichen forming fungi, the family Cladoniaceae in particular,
- Wood rotting fungi, Poorly known Basidiomycetes, Genus Anthracoidea
- Petrology and mineralogy in Earth Sciences
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