A revision of Ceratocarcinus White, 1847, and Harrovia Adams & White, 1849 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with crinoids

The crabs of the genera Ceratocarcinus White, 1847, and Harrovia Adams & White, 1849, are revised. Members of both genera are obligate symbionts of crinoids. Ceratocarcinus White, 1847, now contains three species: C. longimanus White, 1847, C. frontodentata (Shen, Dai & Chen 1982), and C. trilobatus (Sakai, 1938). One species, Ceratocarcinus spinosus Miers, 1879, is referred to a new genus, Tiaramedon. Harrovia is redefined and now contains seven species: H. albolineata Adams & White, 1849, H. cognata, new species, H. elegans De Man, 1887, H. japonica Balss, 1921, A longipes Lanchester, 1900, H. ngi Chen & Xu, 1992, and H. tuberculata Haswell, 1880. Harrovia purpureus Gordon, 1934, and Harrovia egeriae Gordon, 1947, are referred to two new genera, Permanotus and Tauropus respectively.